(Also, just to clarify, the two main things the Seekers are said to have done wrong was know how to cure Tranquility but not tell their boss, and not investigate Meredith as religiously as they should have. The latter of these I do not condone, but the former... well, the Tranquility cure, when used on a mage, does not end in a stable mage, and mages absolutely need to be stable given how their power interacts with their emotions. So there is at least one argument for what the Seekers did as far as the Tranquility cure.)
The crime of the Lord Seeker was to feed the other Seekers with Red Lyrium to kill'em all, while allying with Corypheus because he joined that crazy sect that wanted the end of the world. Lord Seeker Lucius even permitted an envy demon to take his identity to pursue his own agenda of destroying the Seeker order for good, leaving the Templar order in the hands of Samson and Corypheus. And, unlike the Wardens that were mind-controlled or deceived by a false Calling, Lucius did all those stuff on his free will (yet, nobody blames the Seekers for actually working with the enemy on their own free will, instead of the other groups, that were deceived or manipulated to do so).
Then, there is the Tranquility cure and other stuff that Cassandra discovered while in her personal quest.





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