Again, they did not alter the keepers, they altered the signal (see my last post).
Not sure why some people say otherwise - this is clearly stated in the first game.
I do agree that the keeper situation is just as convoluted though. Somehow the reapers built a station that had these maintenance workers and then they built it so that the keepers activate the relay when sent a signal though the station from the reapers. It's a bit of a mess. But at least in ME1 it is acknowledged that it's a convoluted setup. Vigil does speculate that the keepers evolved over time, which changed them in a way so that the reapers could no longer signal them directly. At least it's clear the writers thought about it back in the day.
The fact that the map of the Citadel Tower in ME1 is an outline of a Reaper indicates the writers did envision tying something back to the Citadel.





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