From the Wiki.The Rachni Queen on Noveria mentions "a tone from space" that forced the
rachni to "resonate with its own sour yellow note". This hints that
there may have been more to the Rachni Wars than the Council was aware
of. This is confirmed in Mass Effect 2, depending on whether you
released the Queen, by a message given to Shepard by an asari who is in
service to the newly freed Queen. The asari tells Shepard that the Queen
believes Shepard is fighting the enemy who "soured" the rachni. She
also states that the queen is building an army to fight them when they
arrive in force, based on an otherwise unexplored planet. Shepard
concludes that the message is referring to the Reapers, either through
agents, such as the Collectors, or directly.
Okay, so the Rachni Wars were caused by indoctrination, right? Well, indoctrination by who? Sovereign is a possibility, but his function was to open the Citadel relay, not indoctrinate an entire species (also raises the question of 'where exactly was Sovereign, before the game started?' Did the Reapers send him out 'on foot' once the Citadel Relay didn't open? Was he waiting in the galaxy? Maybe at the Collector Base?).
Any thoughts? If it wasn't Sovereign, then there's another Reaper/Reaper agent floating around the rachni homeworld. I don't think the Collectors can indoctrinate...





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